Note: My ResearcherID is F-9035-2013, see

(for a pdf of any of my papers, please contact me at


Beauchamp, A.T., C.G. Guglielmo, Y.E. Morbey. 2020. Stopover refueling, movement, and departure decisions in the White-throated Sparrow: the influence of intrinsic and extrinsic factors during spring migration. J. Anim. Ecol. Accepted.

Senner, N.R., Y.E. Morbey, B.K. Sandercock. 2020. Editorial: Flexibility in the migration strategies of animals. Front. Ecol. Evol. 8:111. doi:10.3389/fevo.2020.00111

Morbey, Y.E., A. Hedenström. 2020. Leave earlier or travel faster? Optimal mechanisms for managing arrival time in migratory songbirds. Front. Ecol. Evol. 7:492. doi:10.3389/fevo.2019.00492


Deakin, J.E., C.G. Guglielmo, Y.E. Morbey. 2019 Sex differences in spring migratory restlessness in a Nearctic-Neotropical songbird. Auk. 136:1-13. doi:10.1093/auk/ukz017

Thorn, M.W., Y.E. Morbey. 2019. Survival under the gravel: the effects of egg size and parental identity on the in-situ hatching success of Chinook Salmon. Trans. Am. Fish. Soc. 148:991-1003. doi:10.1002/tafs.10192

Seewagen, C.L., Y. Ma, Y.E. Morbey, C.G. Guglielmo. 2019. Stopover departure behavior and flight orientation of spring-migrant Yellow-rumped Warblers (Setophaga coronata) experimentally exposed to methylmercury. J. Ornithol. 160:617-624. doi: 10.1007/s10336-019-01641-2

Crewe, T.L.*, J.E. Deakin, A.T. Beauchamp, Y.E. Morbey. 2019. Detection range of songbirds using a stopover site by automated radio-telemetry. J. Field Ornithol. 90:176-189.

Thorn, M.W., M.F. Dick, L. Oviedo, C.G. Guglielmo, and Y.E. Morbey. 2019. Transgenerational effects of egg nutrients on the early development of Chinook salmon (Oncorhynchus tshawytscha) across a thermal gradient. Can. J. Fish. Aquat. Sci. 76:1253-1262. doi:10.1139/cjfas-2018-0013


Morbey, Y.E. 2018. Female-biased sexual dimorphism in size and age at maturation is reduced at high latitudes in lake whitefish Coregonus clupeaformis. J. Fish Biol. 93:40-46. doi:10.1111/jfb.13675

Morbey, Y.E., and M. Mema. 2018. Size-selective fishing and the potential for fisheries-induced evolution in lake whitefish. Evol. Appl. 11:1412-1424. doi:10.1111/eva.12635.

Hays, Q.R., Y.E. Morbey, K.A. Hobson, J.E. Lyons, and C.G. Guglielmo. 2018. Effects of migration distance and sex on stopover timing and refueling by Wilson’s Warblers. J. Field Ornithol. 89:149-164. doi:10.1111/jofo.12246.

Morbey, Y.E., C.G. Guglielmo, P. Taylor, I. Maggini, J. Deakin, S. Mackenzie. J.M. Brown, and L. Zhao. 2018. Evaluation of sex differences in the stopover behavior and postdeparture movements of wood-warblers. Behav. Ecol. 29:117-127. doi: 10.1093/beheco/arx123.

Thorn, M.W., and Y.E. Morbey. 2018. Egg size and the adaptive capacity of early life history traits in Chinook salmon (Oncorhynchus tshawytscha). Evol. Appl. 11:205-219. doi: 10.1111/eva.12531


Zhao, Y., and Y.E. Morbey. 2017. Estimating the size selectivity of trap nets using a gillnet selectivity experiment: method development and application to lake whitefish in Lake Huron. N. Am. J. Fish. Manage. 37:1341–1349. doi: 10.1080/02755947.2017.1381206

Marklevitz, S.A.C., and Y.E. Morbey. 2017. Habitat use and arrival timing of hatchery and naturalized Chinook salmon spawning in a Great Lakes tributary. Trans. Am. Fish. Soc. 146:567-583. doi:10.1080/00028487.2017.1300606

Kennedy, L.V., Y.E. Morbey, S.A. Mackenzie, P.D. Taylor, and C.G. Guglielmo. 2017. A field test of the effects of body composition analysis by quantitative magnetic resonance of body composition analysis on songbird stopover behaviour. J. Ornithol. 158:593-601. doi:10.1007/s10336-016-1399-2


Gerson, M.A., S.A.C. Marklevitz and Y.E. Morbey. 2016. Timing of spawning and predicted fry emergence by naturalized Chinook salmon (Oncorhynchus tshawytscha) in a Lake Huron tributary. J. Great Lakes Res. 42:678-686. doi:10.1016/j.jglr.2016.03.008

Marklevitz, S.A.C., B.J. Fryer, J. Johnson, D. Gonder, and Y.E. Morbey. 2016. Otolith microchemistry reveals spatio-temporal heterogeneity of natal sources and inter-basin migrations of Chinook salmon in Lake Huron. J Great Lakes Res. 42:668-677. doi: 10.1016/j.jglr.2016.03.007

Bazzi, G., A. Galimberti, Q.R. Hays, I. Bruni, J.G. Cecere, L. Gianfranceschi, KA. Hobson, Y.E. Morbey, N. Saino, C.G. Guglielmo, D. Rubolini. 2016. Adcyap1 polymorphism covaries with breeding latitude in a Nearctic migratory songbird, the Wilson’s warbler (Cardellina pusilla). Ecol. Evol. 6:3226-3239. doi: 10.1002/ece3.2053

Schmaljohann, H., C. Meier, D. Arlt, F. Bairlein, H. van Oosten, Y.E. Morbey, S. Åkesson, M. Buchmann, N. Chernetsov, R. Desaever, J. Elliott, M. Hellström, F. Liechti, A. López, J. Middleton, U. Ottosson, T. Pärt, F. Spina. 2016. Proximate causes of avian protandry differ between subspecies with contrasting migration challenges. Behav. Ecol. 27:321-331. doi:10.1093/beheco/arv160


Thorn, M.W., and Y.E. Morbey. 2015. Evidence for the secondary sexual development of the anal fin in female kokanee salmon Oncorhynchus nerka. J. Fish Biol. doi:10.1111/jfb.12800.


Morbey, Y.E., E.L. Jensen, and M. Russello. 2014. Time scale matters: genetic analysis does not support adaptation-by-time as the mechanism for adaptive seasonal declines in kokanee reproductive lifespan. Ecol. Evol. doi: 10.1002/ece3.1214.


Morbey, Y.E., and B.J. Shuter. 2013. Intermittent breeding in the absence of a large cost of reproduction: evidence for a non-migratory, iteroparous salmonid. Ecosphere 4(12):150 (9 pp). doi:10.1890/ES13-00259.

Morbey, Y.E. 2013. Protandry, sexual size dimorphism, and adaptive growth. J. Theor. Biol. 339:93-99. doi: 10.1016/j.jtbi.2013.05.009.

Mizzau, T.W., S.R. Garner, S.A.C. Marklevitz, G.J. Thompson, and Y.E. Morbey. 2013. A genetic test of sexual size dimorphism in pre-emergent Chinook salmon. PLoS One 8:E78421.

Seewagen, C.L., C.G. Guglielmo, and Y.E. Morbey. 2013. Stopover refueling rate underlies protandry and seasonal variation in migration timing of songbirds. Behav. Ecol. 24:634-642. doi:10.1093/beheco/ars225.


Warren, M.A., and Y.E. Morbey, Y.E. 2012. Migration timing in female kokanee salmon: diel patterns and effects of maturation state. J. Fish Biol. 81:1234-1247. doi:10.1111/j.1095-8649.2012.03402.x

Morbey, Y.E., T. Coppack, F. Pulido. 2012. Adaptive hypotheses for protandry in arrival to breeding areas: a review of models and empirical tests. J. Ornithol. 153(Suppl. 1):207-215. DOI:10.1007/s10336-012-0854-y.

Suk, H.Y., B.D. Neff, K. Quach, and Y.E. Morbey. 2012. Evolution of introduced Chinook salmon (Oncorhynchus tshawytscha) in Lake Huron: emergence of population genetic structure in less than 10 generations. Ecol. Freshw. Fish 21:235-244.


Warren, M.A., and Y.E. Morbey. 2011. Reproductive timing phenotypes in female salmon: true alternatives or extreme variants? Anim. Behav. 82:1373-1380.

Marklevitz, S.A.C., B.J. Fryer, D. Gonder, Z. Yang, J. Johnson, A. Moerke, and Y.E. Morbey. 2011. Use of otolith chemistry to discriminate juvenile Chinook salmon (Oncorhynchus tshawytscha) from different wild populations and hatcheries in Lake Huron. J. Great Lakes Res. 37:698-706

McLeod, D.V., R.A. Cottrill, and Y.E. Morbey. 2011. Sea lamprey wounding in Canadian waters of Lake Huron from 2000 to 2009: temporal changes differ among regions. J. Great Lakes Res. 37:601-608.

Earlier than 2010

Morbey, Y.E., P. Couture, P. Busby, and B.J. Shuter. 2010. Physiological correlates of seasonal growth patterns in lake trout Salvelinus namaycush. J. Fish Biol. 77:2298-2314.

Jastrebski, C.J. and Y.E. Morbey. 2009. Egg size variation in lake trout: phenotype-habitat correlations show an effect of rearing environment. Trans. Am. Fish. Soc. 138:1342-1351.

Morbey, Y.E., D.M. Anderson, and B.A. Henderson. 2008. Progress towards the rehabilitation of lake trout (Salvelinus namaycush) in South Bay, Lake Huron. J. Great Lakes Res. 34:287-300.

Morbey, Y.E., and A.P. Hendry. 2008. Adaptation to spawning habitat in salmonids. In Salmon spawning habitat in rivers: physical controls, biological responses, and approaches to remediation. Edited by D. Sear and P. DeVries. American Fisheries Society, Symposium 65, Bethesda, Maryland. pp. 15-36.

Morbey, Y.E., K. Vascotto, and B.J. Shuter. 2007. Dynamics of piscivory in lake trout following a smallmouth bass invasion: a historical reconstruction. Trans. Am. Fish. Soc. 136:477-483.

Morbey, Y.E., P. Addison, B.J. Shuter, and K. Vascotto. 2006. Within-population heterogeneity of habitat use by lake trout Salvelinus namaycush. J. Fish Biol. 69:1675-1696.

Morbey, Y.E., and C.G. Guglielmo. 2006. Evidence of a novel reproductive tactic in female kokanee Oncorhynchus nerka. J. Fish Biol. 69:1731-1743.

Morbey, Y.E., A.P. Hendry, and C.E. Brassil. 2005. Rapid senescence in Pacific salmon. Am. Nat. 166:556-568.

Morbey, Y.E., and P.A. Abrams. 2004. The interaction between reproductive lifespan and protandry in seasonal breeders. J Evol. Biol. 17:768-778.

Hendry, A.P., Y.E. Morbey, O.K. Berg, and J.K. Wenburg. 2004. Adaptive variation in senescence: reproductive life span in a wild salmon population. Proc Roy Soc Lond B 271:259-266.

Morbey, Y.E., and R.C. Ydenberg. 2003. Timing games in the reproductive phenology of female Pacific salmon (Oncorhynchus spp.). Am Nat 161:284-298.

Morbey, Y.E. 2003. Pair formation, pre-spawning waiting, and protandry in kokanee Oncorhynchus nerka. Behav Ecol Sociobiol 54:127-135.

Morbey, Y.E. 2002. Protandry models and their application to salmon. Behav Ecol 13:337-343.

Morbey, Y.E. 2002. The mate-guarding behaviour of male kokanee (Oncorhynchus nerka). Behaviour 139:507-528.

Morbey, Y.E., and R.C. Ydenberg. 2001. Protandrous arrival timing to breeding areas: a review. Ecol Lett 4:663-673.

Morbey, Y. 2000. Protandry in Pacific salmon. Can J Fish Aquat Sci 57:1252-1257.

Morbey, Y.E., and R.C. Ydenberg. 2000. Seasonal decline in nestling growth: support for the parental-quality hypothesis in Cassin's Auklets. Auk 117:1065-1068.

Morbey, Y.E., R.C. Ydenberg, H.A. Knechtel, and A. Harfenist. 1999. Parental provisioning, nestling departure decisions, and prefledging mass recession in Cassin's Auklets. Anim Behav 57:873-881.

Morbey, Y.E., and R.C. Ydenberg. 1997. Intraspecific variability in nestling growth and fledging behavior of Cassin's Auklets at Triangle Island, British Columbia. Condor 99:361-371.

Morbey, Y.E. 1996. The abundance and effects of ticks (Ixodes uriae) on nestling Cassin's Auklets (Ptychoramphus aleuticus) at Triangle Island, British Columbia. Can J Zool 74:1585-1589.